Sly Cooper Mysteries of Thieves is the fifth Sly Cooper game, preceded by Sly Cooper Thieves in Time.
After the events of Sly 4: Thieves in Time, Bentley, on the search for Sly Cooper who went missing since the destruction on Le Paradox's blimp, notices a mysterious variety of signs marking the getaway heists of the infamous Revolution Gang on their attempts to wipe out a family thieving clan for a mysterious ghost which looked similar to the Cooper Clan nemesis; Clockwerk. Intrigued of this Revolution business, Bentley rallies up the entire Cooper gang to travel through time in search of Sly Cooper and to solve all the mysteries of whoever and whatever is about to destroy the Cooper clan history where they'll face the ultimate heists and hazardous getaways along the way.
The main protagonist of the series, lost in time since the Le Paradox blimp destruction. Same details from the previous game.
He is both in his attire from Sly 3: Honor Among Thieves with his wheelchair upgraded.
Murray was in the fighting championship which reached into great heights of success.
Carmelita returns as Interpol's most successful inspector with a new shock pistol that can lock on like the old one
she used from Sly 4: Thieves in Time.
Dimitri returns as the fully playable character with his diving equipment from Sly 3: Honor Among Thieves and also a baggage of disguises he'll be using for certain jobs.
Seen from Sly 4: Thieves in Time on the bad side, but later revealed that the real Penelope was abducted by Le Paradox, exiled into the era of WW1 while her evil twin sister Emma took her place in disguise of Penelope herself. But the real Penelope will return as another playable character, recruited back to the Cooper gang.
The outback mystic for the gang, returning as the fully playable character with new feats of skills unlike last time in Sly 3: Honor Among Thieves.
A former cosmetic-owning Monitor Lizard from present-day New York City, brought to Ancient Egypt in order to swipe Slytunkhamen's canes and replace him with her dictator plans to terrorize Egypt with her cosmetic brands.
A former fisherman Seal from Eastern Wales who was involved in illegal whale-hunting since the recent sperm whale attack on his leg, sent to handed over the authorities of the time period by Henrietta "One-Eye" Cooper whose dagger hook he'd been ordered to steal.
A former Mongolian general from a sinister line of thieves, brought to Celtic Scotland to swipe Slaigh's cane and terrorize Scotland with a army of undead creatures (stolen prehistoric skeletons and paleo statues from museums).
The Red Dragon
The Red Dragon is actually Penelope's evil twin Emma in disguise, ordered to kill Otto Van Cooper and sabotage his mechanical machinery.
Hogan the Tyrant
A cigar-smoking rhinoceros from present-day Finland, brought to the Victorian London era to frame Thaddeus Winslow Cooper and swipe her cane away.
Escaped from prison, brought into the 1940's in Chicago, ordered to arrest Curtis Capone Cooper whose gatling gun-cane he'd been sent to steal, but later betrayed and crushed to death by Clockwerk.
A ancient nemesis of the Cooper clan, thought to defeated by Sly, but his ghost will appear as the main antagonist in his attempt to wipe out the Cooper clan like before, this time with vengeance.
Ancestors Featured In-Game
Sly's Egyptian ancestor with the ability to turn invisible to escape the greedy nobleman.
Henrietta One Eye Cooper
The famous female pirate from the 18th century, thieving numerous bands of pirates from coast to coast in the Caribbean Seas.
The stongest Cooper from Celtic Scotland with two occupations: a warmonger and a blacksmith.
Thaddeus Winslow Cooper
The most chivalrous gentlemanly Cooper in secret service with Scotland Yard.
Otto Van Cooper
The mechanic from WW1 in Germany, supplying formidable machinery for the war.
Curtis Capone Cooper
Sly's great grandfather from the 1940's, also given the reputation as the most enforcing gangster of Chicago.